東北棕熊

東北棕熊
保护状况
	; 易危（IUCN 2.3）[1]
科学分类
界：	动物界 Animalia
门：	脊索动物门 Chordata
纲：	哺乳纲 Mammalia
目：	食肉目 Carnivora
科：	熊科 Ursidae
属：	熊属 Ursus
种：	棕熊 U. arctos
亚种：	東北棕熊 U. a. lasiotus
三名法
	Ursus arctos lasiotus; Gray, 1867[2]
異名
	baikalensis Ognev, 1924; cavifrons (Heude, 1901); ferox Temminck, 1844; macneilli Lydekker, 1909; melanarctos Heude, 1898; yesoensis Lydekker, 1897

東北棕熊（學名：Ursus arctos lasiotus），又名烏蘇里棕熊、东北马熊，是棕熊的一個亞種，分佈于西伯利亞、朝鮮半島、中國東北和北海道，與堪察加棕熊相似，但毛色更黑、頭骨更長。其分佈于北海道的種群有時會單獨列為一個亞種，即日本棕熊（Ursus arctos yesoensis）。它是體型較大的棕熊之一，最大的東北棕熊個頭體重超過400公斤，堪比阿拉斯加棕熊。[3]

習性

頭骨

東北棕熊食譜很雜，它以各類植物漿果、根莖、種子和小型、大型動物為食，甚至也吃螞蟻[4]。

東北虎有時會捕食東北棕熊，但這種情況很少出現[5][6]。此外，東北虎捕食的一般是幼熊，但也有東北虎襲擊冬眠的成年雌性東北棕熊的記錄[7][8][9]。1944-1959年，俄羅斯遠東地區共錄下32起東北虎襲擊東北棕熊的案例[10][11][12]。

1915年12月，日本北海道發生三毛別棕熊襲擊事件，造成7人死亡、3人重傷，為日本史上最嚴重的野生動物攻擊事件。

參考文獻

McLellan, B.N.; Servheen, C. & Huber, D. . IUCN Red List of Threatened Species. 2008. [27 January 2009].
Gray, J. E. . Annals and Magazine of Natural History. Ser. 3. 1867, 20 (118): 301. doi:10.1080/00222936708694139.
Mammals of the Soviet Union Vol.II Part 1a, SIRENIA AND CARNIVORA (Sea cows; Wolves and Bears), V.G Heptner and N.P Naumov editors, Science Publishers, Inc. USA. 1998. ISBN 1-886106-81-9
Onoyama, Keiichi. (PDF). Res. Bull. Obihiro University (Obihiro University of Agriculture and Veterinary Medicine). 1987-11-16, 15: 313–318 [2009-05-05]. （原始内容 (PDF)存档于2011-07-22）.
Seryodkin; 等. . Tigers of Sikhote-Alin Zapovednik: Ecology and Conservation. 2005: 156–163.
Geptner, V. G., Sludskii, A. A. (1972). Mlekopitaiuščie Sovetskogo Soiuza. Vysšaia Škola, Moskva. (In Russian; English translation: Heptner, V. G.; Sludskii, A. A.; Bannikov, A. G.; (1992). the Soviet Union. Volume II, Part 2: Carnivora (Hyaenas and Cats). Smithsonian Institution and the National Science Foundation, Washington DC). Pp. 95–202.
Frasef, A. . CABI. 2012: 72–77. ISBN 978-1-84593-926-7.
Prynn, David. . Russian Nature Press. 2004: 115.
V.G. Heptner and N.P. Naumov. Mammals of the Soviet Union. Vol 2 IIa 1998. New Delhi, India: Amerind Publishing. p671
Mammals of the Soviet Union Volume 2, by V.G Heptner & A.A. Sludskii, p177
Seryodkin, I. V., J. M. Goodrich, A. V. Kostyrya, B. O. Schleyer, E. N. Smirnov, L. L. Kerley, and D. G. Miquelle (2005). "Relationship between tigers, brown bears, and Himalayan black bears". pp. 95-202 in D. G. Miquelle, E. N. Smirnov, and J. M. Goodrich (eds.), Tigers of Sikhote-Alin Zapovednik: Ecology and Conservation. Vladivostok, Russia: PSP.
Table 1 页面存档备份，存于. Location, physical status, size and circumstances of deaths of Amur tiger males in the Russian Far East, 1970-1994.

外部連接

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[1] McLellan, B.N.; Servheen, C. & Huber, D. . IUCN Red List of Threatened Species. 2008. [27 January 2009].

[2] Gray, J. E. . Annals and Magazine of Natural History. Ser. 3. 1867, 20 (118): 301. doi:10.1080/00222936708694139.

[soviet-3] Mammals of the Soviet Union Vol.II Part 1a, SIRENIA AND CARNIVORA (Sea cows; Wolves and Bears), V.G Heptner and N.P Naumov editors, Science Publishers, Inc. USA. 1998. ISBN 1-886106-81-9

[ants-4] Onoyama, Keiichi. (PDF). Res. Bull. Obihiro University (Obihiro University of Agriculture and Veterinary Medicine). 1987-11-16, 15: 313–318 [2009-05-05]. （原始内容 (PDF)存档于2011-07-22）.

[Seryodkin2-5] Seryodkin; 等. . Tigers of Sikhote-Alin Zapovednik: Ecology and Conservation. 2005: 156–163.

[Geptner1972-6] Geptner, V. G., Sludskii, A. A. (1972). Mlekopitaiuščie Sovetskogo Soiuza. Vysšaia Škola, Moskva. (In Russian; English translation: Heptner, V. G.; Sludskii, A. A.; Bannikov, A. G.; (1992). the Soviet Union. Volume II, Part 2: Carnivora (Hyaenas and Cats). Smithsonian Institution and the National Science Foundation, Washington DC). Pp. 95–202.

[FBW-7] Frasef, A. . CABI. 2012: 72–77. ISBN 978-1-84593-926-7.

[amur-8] Prynn, David. . Russian Nature Press. 2004: 115.

[9] V.G. Heptner and N.P. Naumov. Mammals of the Soviet Union. Vol 2 IIa 1998. New Delhi, India: Amerind Publishing. p671

[10] Mammals of the Soviet Union Volume 2, by V.G Heptner & A.A. Sludskii, p177

[11] Seryodkin, I. V., J. M. Goodrich, A. V. Kostyrya, B. O. Schleyer, E. N. Smirnov, L. L. Kerley, and D. G. Miquelle (2005). "Relationship between tigers, brown bears, and Himalayan black bears". pp. 95-202 in D. G. Miquelle, E. N. Smirnov, and J. M. Goodrich (eds.), Tigers of Sikhote-Alin Zapovednik: Ecology and Conservation. Vladivostok, Russia: PSP.

[12] Table 1 页面存档备份，存于. Location, physical status, size and circumstances of deaths of Amur tiger males in the Russian Far East, 1970-1994.