異蟲綱
异虫纲(学名:Xenusia),是叶足动物的代表演化支,可能包括有爪动物祖先。[2]模式属是异虫属。[3]它们具有相对较大的环状圆柱形的身体,它们的叶足基部有结节。有些附肢较大,[3]这些埋藏的化石信息能够做为其代表。[4]它们的口器在躯体末端或接近末端,它们曾经生活在全球各地的海洋当中。[5]它们可能是一个並系群的代表,而不是单系群。
异虫纲 化石时期:寒武纪第三期–早石炭世 | |
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广卫怪诞虫 Hallucigenia hongmeia | |
科学分类 | |
界: | 动物界 Animalia |
亚界: | 真后生动物亚界 Eumetazoa |
演化支: | 副同源异形基因动物 ParaHoxozoa |
演化支: | 浮浪幼虫样动物 Planulozoa |
演化支: | 两侧对称动物 Bilateria |
演化支: | 肾管动物 Nephrozoa |
演化支: | 原口动物 Protostomia |
总门: | 蜕皮动物总门 Ecdysozoa |
演化支: | 泛节肢动物 Panarthropoda |
并系群: | †葉足動物 Lobopodia |
纲: | †異蟲綱 Xenusia Dzik & Krumbiegel, 1989[1] |
下属分类 | |
见内文 |
分类
始有爪目 Archonychophora
- 始有爪目 Archonychophora Hou & Bergstrom, 1995[2]– undifferentiated appendages; each segment the same as each other.[6]
- 啰哩山虫科 Luolishaniidae Hou & Bergstrom, 1995 – three small rounded sclerites per segment – these are thorn-shaped spines.[6]
- 啰哩山虫属 Luolishania Hou & Chen 1989
- 神奇啰哩山虫属 Miraluolishania Liu & Shu 2004
- 贫腿虫科 Paucipodiidae Hou et al., 2004[7]– with no dorsal sclerites, two claws, fine annulation and few segments.
- 贫腿虫属 Paucipodia Hou et al. 2004
- 啰哩山虫科 Luolishaniidae Hou & Bergstrom, 1995 – three small rounded sclerites per segment – these are thorn-shaped spines.[6]
These taxa are only known from the Chengjiang (Cambrian Stage 2 Series 3).
原有爪目 Protonychophora
- 原有爪目 Protonychophora Hutchinson, 1930 – spiny legs, reduced tail.
- 埃谢栉蚕科 Aysheaiidae Walcott, 1911 – many claws on each leg. Anterior grasping appendages with long spines.
- 埃谢栉蚕属 Aysheaia Walcott 1911
- 异虫科 Xenusiidae Dzik & Krumbiegel, 1989 – >20 leg-bearing segments; paired, rounded sclerites on each segment; spiny legs.
- 异虫属 Xenusion Pompeckj, 1927
- 尖山叶足虫属 Jianshanopodia Liu et al.[3]– from Chengjiang; two rows of tubercles but no obvious sclerites.
- Hadranax Budd & Peel 1998 – from the Sirius Passet. Lacks obvious dorsal armature, but bears four 'nodes' per row.
- 埃谢栉蚕科 Aysheaiidae Walcott, 1911 – many claws on each leg. Anterior grasping appendages with long spines.
坚有爪目 Scleronychophora
- 坚有爪目 Scleronychophora Hou & Bergstrom, 1995 – paired sclerites; elongated head sclerites.
- 始小贝科 Eoconchariidae Hou & Shu, 1987 – ~10 leg-bearing segments, small head, sieve-like sclerites, varying in shape along body; curved claws on annulated legs; terminal mouth and anus.
- 微网虫属 Microdictyon Bengtson et al.
- Quadratapora Hao and Shu, 1987
- Fusuconcharium Hao and Shu, 1987
- 怪诞虫科 Hallucigeniidae Conway Morris, 1977 – ~10 leg-bearing segments; large sclerites covering head; sclerites on each segment are long spines.
- 怪诞虫属 Hallucigenia Conway Morris 1977 – from the Chengjiang, Kaili and Burgess Shale.[8]
- 心网虫科 Cardiodictyidae Hou & Bergstrom, 1995 – many segments (~23); large sclerites cover head; hexagonal sclerite on each body segment.
- 心网虫属 Cardiodictyon Hou et al. 1991 – known from the Chengjiang.[8]
- 始小贝科 Eoconchariidae Hou & Shu, 1987 – ~10 leg-bearing segments, small head, sieve-like sclerites, varying in shape along body; curved claws on annulated legs; terminal mouth and anus.
等有爪目 Paronychophora
Order unassigned
'Phylum' Onychophora
Onychophora are distinguished by their terrestrial habit, their ventral mouth; their antennae, jaws and oral papillae; they seem to be most closely related to the Paronychophora. Their first fossils are Carboniferous (Helenodora), although they may have had a cryptic earlier history.
參考文獻
- Dzik, J.; Krumbiegel, G. N. . Lethaia. 1989, 22 (2): 169–181. doi:10.1111/j.1502-3931.1989.tb01679.x.
- Hou, X.; Bergström, J. A. N. . Zoological Journal of the Linnean Society. 1995, 114: 3–19. doi:10.1111/j.1096-3642.1995.tb00110.x.
- Jianni Liu Degan Shu, Jian Han, Zhifei Zhang & Xingliang Zhang. (PDF). Acta Palaeontologica Polonica. 2006, 51 (2): 215–222 [2020-04-27]. (原始内容存档 (PDF)于2017-08-09).
- Julián Monge-Nájera & Xianguang Hou; Hou. (PDF). Revista de Biología Tropical. 2002, 50 (3–4): 1133–1138. PMID 12947596. (原始内容 (PDF)存档于June 29, 2011). 已忽略未知参数
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(帮助) - George Poinar, Jr. . Invertebrate Biology. 2000, 119 (1): 104–109. JSTOR 3227105. doi:10.1111/j.1744-7410.2000.tb00178.x.
- Ma, X.; Hou, X.; Bergström, J. . Arthropod Structure & Development. 2009, 38 (4): 271–291. doi:10.1016/j.asd.2009.03.001.
- Xian-Guang Hou, Xiao-Ya Ma, Jie Zhao & Jan Bergström; Ma; Zhao; Bergström. . Lethaia. 2004, 37 (3): 235–244. doi:10.1080/00241160410006555.
- Whittle, R. J.; Gabbott, S. E.; Aldridge, R. J.; Theron, J. . Palaeontology. May 2009, 52 (3): 561–567. doi:10.1111/j.1475-4983.2009.00860.x.
- Liu, J.; Shu, D.; Han, J.; Zhang, Z.; Zhang, X. . Acta Palaeontologica Polonica. 2008, 53 (2): 285–292. doi:10.4202/app.2008.0209. 已忽略未知参数
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(帮助) - Maas, A.; Mayer, G.; Kristensen, R. M.; Waloszek, D. . Chinese Science Bulletin. 2007, 52 (24): 3385–3392. doi:10.1007/s11434-007-0515-3.
- Haug, J. T.; Mayer, G.; Haug, C.; Briggs, D. E. G. . Current Biology. 2012, 22 (18): 1673–1675. PMID 22885062. doi:10.1016/j.cub.2012.06.066. 已忽略未知参数
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(帮助) - Gámez Vintaned, J. A.; Liñán, E.; Zhuravlev, A. . . 29 June 2011: 193–219. ISBN 978-3-642-20762-4. doi:10.1007/978-3-642-20763-1_12.
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