異蟲綱

异虫纲(学名:Xenusia),是叶足动物的代表演化支,可能包括有爪动物祖先。[2]模式属是异虫属[3]它们具有相对较大的环状圆柱形的身体,它们的叶足基部有结节。有些附肢较大,[3]这些埋藏的化石信息能够做为其代表。[4]它们的口器在躯体末端或接近末端,它们曾经生活在全球各地的海洋当中。[5]它们可能是一个並系群的代表,而不是单系群。

异虫纲
化石时期:寒武纪第三期早石炭世
PreЄ
广卫怪诞虫 Hallucigenia hongmeia
科学分类
界: 动物界 Animalia
亚界: 真后生动物亚界 Eumetazoa
演化支 副同源异形基因动物 ParaHoxozoa
演化支 浮浪幼虫样动物 Planulozoa
演化支 两侧对称动物 Bilateria
演化支 肾管动物 Nephrozoa
演化支 原口动物 Protostomia
总门: 蜕皮动物总门 Ecdysozoa
演化支 泛节肢动物 Panarthropoda
并系群 葉足動物 Lobopodia
纲: 異蟲綱 Xenusia
Dzik & Krumbiegel, 1989[1]
下属分类

见内文

分类

异虫纲包括以下:[5]

始有爪目 Archonychophora

  • 始有爪目 Archonychophora Hou & Bergstrom, 1995[2]– undifferentiated appendages; each segment the same as each other.[6]
    • 啰哩山虫科 Luolishaniidae Hou & Bergstrom, 1995 – three small rounded sclerites per segment – these are thorn-shaped spines.[6]
      • 啰哩山虫属 Luolishania Hou & Chen 1989
      • 神奇啰哩山虫属 Miraluolishania Liu & Shu 2004
    • 贫腿虫科 Paucipodiidae Hou et al., 2004[7]– with no dorsal sclerites, two claws, fine annulation and few segments.
      • 贫腿虫属 Paucipodia Hou et al. 2004

These taxa are only known from the Chengjiang (Cambrian Stage 2 Series 3).

原有爪目 Protonychophora

  • 原有爪目 Protonychophora Hutchinson, 1930 – spiny legs, reduced tail.
    • 埃谢栉蚕科 Aysheaiidae Walcott, 1911 – many claws on each leg. Anterior grasping appendages with long spines.
      • 埃谢栉蚕属 Aysheaia Walcott 1911
    • 异虫科 Xenusiidae Dzik & Krumbiegel, 1989 – >20 leg-bearing segments; paired, rounded sclerites on each segment; spiny legs.
      • 异虫属 Xenusion Pompeckj, 1927
      • 尖山叶足虫属 Jianshanopodia Liu et al.[3]– from Chengjiang; two rows of tubercles but no obvious sclerites.
      • Hadranax Budd & Peel 1998 – from the Sirius Passet. Lacks obvious dorsal armature, but bears four 'nodes' per row.

坚有爪目 Scleronychophora

  • 坚有爪目 Scleronychophora Hou & Bergstrom, 1995 – paired sclerites; elongated head sclerites.
    • 始小贝科 Eoconchariidae Hou & Shu, 1987 – ~10 leg-bearing segments, small head, sieve-like sclerites, varying in shape along body; curved claws on annulated legs; terminal mouth and anus.
      • 微网虫属 Microdictyon Bengtson et al.
      • Quadratapora Hao and Shu, 1987
      • Fusuconcharium Hao and Shu, 1987
    • 怪诞虫科 Hallucigeniidae Conway Morris, 1977 – ~10 leg-bearing segments; large sclerites covering head; sclerites on each segment are long spines.
      • 怪诞虫属 Hallucigenia Conway Morris 1977 – from the Chengjiang, Kaili and Burgess Shale.[8]
    • 心网虫科 Cardiodictyidae Hou & Bergstrom, 1995 – many segments (~23); large sclerites cover head; hexagonal sclerite on each body segment.
      • 心网虫属 Cardiodictyon Hou et al. 1991 – known from the Chengjiang.[8]

等有爪目 Paronychophora

  • 等有爪目 Paronychophora Hou & Bergstrom, 1995 – short, downwards facing head; papillae on body and legs, some arranged in rows; shield-like sclerites on head; dorsal armature of spines;[9]claw-like jaws; annulated legs [Note: diagnosis modified in [6]].

Order unassigned

  • Orstenotubulus Maas et al. 2007 – from the Furongian Orsten deposits – with retractable dorsal spines.[10]
  • Carbotubulus Haug et al. 2012[11]– from the Mazon creek. Dorwal armature uncertain. Few segments, long limbs.
  • Mureropodia[12]from the Stage 2 Murero lagerstatten, Spain.

'Phylum' Onychophora

Onychophora are distinguished by their terrestrial habit, their ventral mouth; their antennae, jaws and oral papillae; they seem to be most closely related to the Paronychophora. Their first fossils are Carboniferous (Helenodora), although they may have had a cryptic earlier history.

參考文獻

  1. Dzik, J.; Krumbiegel, G. N. . Lethaia. 1989, 22 (2): 169–181. doi:10.1111/j.1502-3931.1989.tb01679.x.
  2. Hou, X.; Bergström, J. A. N. . Zoological Journal of the Linnean Society. 1995, 114: 3–19. doi:10.1111/j.1096-3642.1995.tb00110.x.
  3. Jianni Liu Degan Shu, Jian Han, Zhifei Zhang & Xingliang Zhang. (PDF). Acta Palaeontologica Polonica. 2006, 51 (2): 215–222 [2020-04-27]. (原始内容存档 (PDF)于2017-08-09).
  4. Julián Monge-Nájera & Xianguang Hou; Hou. (PDF). Revista de Biología Tropical. 2002, 50 (3–4): 1133–1138. PMID 12947596. (原始内容 (PDF)存档于June 29, 2011). 已忽略未知参数|url-status= (帮助)
  5. George Poinar, Jr. . Invertebrate Biology. 2000, 119 (1): 104–109. JSTOR 3227105. doi:10.1111/j.1744-7410.2000.tb00178.x.
  6. Ma, X.; Hou, X.; Bergström, J. . Arthropod Structure & Development. 2009, 38 (4): 271–291. doi:10.1016/j.asd.2009.03.001.
  7. Xian-Guang Hou, Xiao-Ya Ma, Jie Zhao & Jan Bergström; Ma; Zhao; Bergström. . Lethaia. 2004, 37 (3): 235–244. doi:10.1080/00241160410006555.
  8. Whittle, R. J.; Gabbott, S. E.; Aldridge, R. J.; Theron, J. . Palaeontology. May 2009, 52 (3): 561–567. doi:10.1111/j.1475-4983.2009.00860.x.
  9. Liu, J.; Shu, D.; Han, J.; Zhang, Z.; Zhang, X. . Acta Palaeontologica Polonica. 2008, 53 (2): 285–292. doi:10.4202/app.2008.0209. 已忽略未知参数|doi-access= (帮助)
  10. Maas, A.; Mayer, G.; Kristensen, R. M.; Waloszek, D. . Chinese Science Bulletin. 2007, 52 (24): 3385–3392. doi:10.1007/s11434-007-0515-3.
  11. Haug, J. T.; Mayer, G.; Haug, C.; Briggs, D. E. G. . Current Biology. 2012, 22 (18): 1673–1675. PMID 22885062. doi:10.1016/j.cub.2012.06.066. 已忽略未知参数|doi-access= (帮助)
  12. Gámez Vintaned, J. A.; Liñán, E.; Zhuravlev, A. . . 29 June 2011: 193–219. ISBN 978-3-642-20762-4. doi:10.1007/978-3-642-20763-1_12.
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