肺螺類

有肺類; 化石时期：石炭纪–現在 PreЄ Є O S D C P T J K Pg N
科学分类
界：	动物界 Animalia
门：	軟體動物門 Mollusca
纲：	腹足纲 Gastropoda
演化支：	直腹足類 Orthogastropoda
演化支：	後生腹足類 Apogastropoda
亚纲：	異鰓亞綱 Heterobranchia
–非正式–：	有肺類 Pulmonata; Cuvier, 1814
類群
	基眼類 Basommatophora!; 真有肺總目 Eupulmonata;

肺螺類（學名：），亦作有肺類，舊作有肺目、肺螺目或肺螺亚纲，是腹足綱異鰓類支序之下的一個非正式群組。這個群組包括有螺和蛞蝓，均可以透過其外套膜肺（pallial lung）而不是鰓來呼吸空氣。本群組除了包括有蝸牛和淡水螺，還包括多個海螺的科。

傳統定義的肺螺類分類單元的物種在Jörger et al. (2010)被發現原來是個多系群[1]。肺螺類已知從石炭纪開始就已經開始出現，直到現在[2]。

肺螺類物種有一個心室和一個腎臟，還有一個相互連接、左右對稱的神經系統。外套膜腔位於身體的右側，而且沒有鰓，而是演變成一個佈滿血管的肺。大多數物種都有殼，但沒有口蓋，而且本分類元還包括有幾種沒有外殼的蛞蝓。肺類類物種都雌雄間性，部份群組還擁有用作交配的戀矢（love darts）[3]

1997年分類

在原來的《旁得和林德伯格的腹足類分類 (1997年)》，本支序是一個目級分類元，包括下列各個分類元：

收眼亞目 Systellommatophora Pilsbry, 1948
- - 石磺總科 Onchidioidea Rafinesque, 1815
  - Superfamily Otinoidea H. & A. Adams, 1855
  - 拉索蛞蝓總科 Rathouisioidea Sarasin, 1889
基眼亞目 Basommatophora Keferstein in Bronn, 1864：淡水螺及池塘的螺
- - Superfamily Acroloxoidea Thiele, 1931
  - 两栖螺总科 Amphiboloidea J.E. Gray, 1840
  - Superfamily Chilinoidea H. & A. Adams, 1855
  - Superfamily Glacidorboidea Ponder, 1986
  - 椎實螺總科 Lymnaeoidea Rafinesque, 1815
  - 扁蜷總科 Planorboidea Rafinesque, 1815
  - 松螺總科 Siphonarioidea J.E. Gray, 1840
真有肺亞目 Eupulmonata Haszprunar & Huber, 1990
- Infraorder Acteophila Dall, 1885 (= formerly Archaeopulmonata)
  - Superfamily Melampoidea Stimpson, 1851
- Infraorder Trimusculiformes Minichev & Starobogatov, 1975
  - Superfamily Trimusculoidea Zilch, 1959
- 柄眼下目 Stylommatophora A. Schmidt, 1856
  Shells of pulmonate styllomatophoran snails in a museum collection
  An artistic but scientifically incorrect version of various European land snails and slugs (one species here is not a pulmonate), their food plants and fungi, and a beetle that eats mollusks, bottom right.
  - 直尿道小目 Orthurethra
  - Superfamily Achatinelloidea Gulick, 1873
  - Superfamily Cochlicopoidea Pilsbry, 1900
  - Superfamily Partuloidea Pilsbry, 1900
  - Superfamily Pupilloidea Turton, 1831
  - 弯尿道小目 Sigmurethra
  - Superfamily Acavoidea Pilsbry, 1895
  - 非洲大蝸牛總科 Achatinoidea Swainson, 1840
  - Superfamily Aillyoidea Baker, 1960
  - Superfamily Arionoidea J.E. Gray in Turnton, 1840
  - Superfamily Buliminoidea Clessin, 1879
  - Superfamily Camaenoidea Pilsbry, 1895
  - 烟管蜗牛总科 Clausilioidea Mörch, 1864
  - Superfamily Dyakioidea Gude & Woodward, 1921
  - Superfamily Gastrodontoidea Tryon, 1866
  - Superfamily Helicoidea Rafinesque, 1815
  - Superfamily Helixarionoidea Bourguignat, 1877
  - Superfamily Limacoidea Rafinesque, 1815
  - Superfamily Oleacinoidea H. & A. Adams, 1855
  - Superfamily Orthalicoidea Albers-Martens, 1860
  - Superfamily Plectopylidoidea Moellendorf, 1900
  - Superfamily Polygyroidea Pilsbry, 1894
  - Superfamily Punctoidea Morse, 1864
  - Superfamily Rhytidoidea Pilsbry, 1893
  - Superfamily Sagdidoidera Pilsbry, 1895
  - Superfamily Staffordioidea Thiele, 1931
  - Superfamily Streptaxoidea J.E. Gray, 1806
  - Superfamily Strophocheiloidea Thiele, 1926
  - Superfamily Trigonochlamydoidea Hese, 1882
  - Superfamily Zonitoidea Mörch, 1864
  - ? Superfamily Athoracophoroidea P. Fischer, 1883 = Tracheopulmonata
  - ? Superfamily Succineoidea Beck, 1837 = Heterurethra

2005年分類

Examples of Pulmonata: Achatina fulica top right, Bielzia coerulans top left, Praticolella berlandieriana center right, Megalobulimus oblongus in the center, Euglandina rosea center left, Helix pomatia bottom right & Ashmunella levettei bottom left

根據《布歇特和洛克羅伊的腹足類分類 (2005年)》，Glacidorboidea被搬到異鰓類支序之下的下異鰓類群組，本分類餘下下列各個分類[4]：

有肺類非正式群組

包括基眼類非正式群組及真有肺類支序。

基眼類非正式群組

基眼類（Basommatophora）非正式群組包括喜濕螺類（Hygrophila）支序和下列兩個總科：

两栖螺总科 Amphiboloidea J.E. Gray, 1840
- 两栖螺科 Amphibolidae
松螺總科 Siphonarioidea J.E. Gray, 1840
- 松螺科 Siphonariidae
- † Family Acroreiidae

喜濕螺類

喜濕螺類（Hygrophila）支序包括下列四個總科：

斜顶螺总科 Acroloxoidea Thiele, 1931
- 斜顶螺科 Acroloxidae Thiele, 1931
Superfamily Chilinoidea Dall, 1870
- Family Chilinidae Dall, 1870
- Family Latiidae Hutton, 1882
椎实螺总科（Lymnaeoidea Rafinesque, 1815）
- 椎实螺科（Lymnaeidae Rafinesque, 1815）
扁蜷總科（Planorboidea Rafinesque, 1815）
- 扁蜷科（Planorbidae Rafinesque, 1815）
  - 扁蜷亞科（Planorbinae Rafinesque, 1815）
    - Tribe Ancylini Rafinesque, 1815：從原來的Ancylidae科降格而成
- 膀胱螺科（Physidae Fitzinger, 1833，又名囊螺科）

真有肺類支序

真有肺類（Eupulmonata）支序包括the clades Systellommatophora and Stylommatophora

Superfamily Trimusculoidea
- Family Trimusculidae
Superfamily Otinoidea
- Family Otinidae
- Family Smeagolidae
耳螺總科 Ellobioidea
- 耳螺科 Ellobiidae

收眼類支序

收眼類支序（Systellommatophora = Gymnomorpha）之下有兩總科[5]，分別為：

石磺總科 Onchidioidea Rafinesque, 1815
- 石磺科 Onchidiidae Rafinesque, 1815
皺足蛞蝓總科 Veronicelloidea Gray, 1840
- 皺足蛞蝓科 Veronicellidae Gray, 1840
- 拉索蛞蝓科 Rathouisiidae Heude, 1885

Clade Stylommatophora

Contains the subclades Elasmognatha, Orthurethra and the informal group Sigmurethra

Subclade Elasmognatha

Superfamily Succineoidea
- Family Succineidae
Superfamily Athoracophoroidea
- Family Athoracophoridae

Subclade Orthurethra

帕图螺超科（）
- 帕图螺科（）
小玛瑙螺超科（）
- 小玛瑙螺科（）
槲果螺超科（）
- 槲果螺科（）
- 同纹螺科（）
蛹蜗牛超科（）
- 虹蛹螺科（）
- 粒蛹螺科（）
- （已灭绝）
- 蛹状螺科（）
- 球果螺科（）
- 瓦娄蜗牛科 Valloniidae Morse, 1864
- 牙蛹蝸牛科 Vertiginidae Fitzinger, 1833
艾纳螺超科（）[6]
- 艾纳螺科（）
- 霜纳螺科（ Wenz, 1923）

Informal Group Sigmurethra

Superfamily Clausilioidea
- Family Clausiliidae
- † Family Anadromidae
- † Family Filholiidae
- † Family Palaeostoidae
Superfamily Orthalicoidea
- Family Orthalicidae
- Family Cerionidae
- Family Coelociontidae
- † Family Grangerellidae
- Family Megaspiridae
- Family Placostylidae
- Family Urocoptidae
Superfamily Achatinoidea
- Family Achatinidae
- Family Ferussaciidae
- Family Micractaeonidae
- Family Subulinidae
Superfamily Aillyoidea
- Family Aillyidae
Superfamily Testacelloidea
- Family Testacellidae
- Family Oleacinidae
- Family Spiraxidae
Superfamily Papillodermatoidea
- Family Papillodermatidae
Superfamily Streptaxoidea
- Family Streptaxidae
Superfamily Rhytidoidea
- Family Rhytididae
- Family Chlamydephoridae
- Family Haplotrematidae
- Family Scolodontidae
Superfamily Acavoidea
- Family Acavidae
- Family Caryodidae
- Family Dorcasiidae
- Family Macrocyclidae
- Family Megomphicidae
- Family Strophocheilidae
Superfamily Punctoidea
- Family Punctidae
- † Family Anastomopsidae
- Family Charopidae
- Family Cystopeltidae
- Family Discidae
- Family Endodontidae
- Family Helicodiscidae
- Family Oreohelicidae
- Family Thyrophorellidae
Superfamily Sagdoidea
- Family Sagdidae

limacoid clade

Superfamily Staffordioidea
- Family Staffordiidae
Superfamily Dyakioidea
- Family Dyakiidae
Superfamily Gastrodontoidea
- Family Gastrodontidae
- Family Chronidae
- Family Euconulidae
- Family Oxychilidae
- Family Pristilomatidae
- Family Trochomorphidae
- Fossil taxa probably belonging to the Gastrodontoidea
  - Subfamily † Archaeozonitinae
  - Subfamily † Grandipatulinae
  - Subfamily † Palaeoxestininae
Superfamily Parmacelloidea
- Family Parmacellidae
- Family Milacidae
- Family Trigonochlamydidae
Superfamily Zonitoidea
- Family Zonitidae
Superfamily Helicarionoidea
- Family Helicarionidae
- Family Ariophantidae
- Family Urocyclidae
Superfamily Limacoidea
- Family Limacidae
- Family Agriolimacidae
- Family Boettgerillidae
- Family Vitrinidae

other Sigmurethra

Two superfamilies belongs to clade Sigmurethra, but they are not in the limacoid clade.

Superfamily Arionoidea
- Family Arionidae
- Family Anadenidae
- Family Ariolimacidae
- Family Binneyidae
- Family Oopeltidae
- Family Philomycidae
Superfamily Helicoidea
- Family Helicidae
- Family Bradybaenidae
- Family Camaenidae
- Family Cepolidae
- Family Cochlicellidae
- Family Elonidae
- Family Epiphragmophoridae
- Family Halolimnohelicidae
- Family Helicodontidae
- Family Helminthoglyptidae
- Family Humboldtianidae
- Family Hygromiidae
- Family Monadeniidae
- Family Pleurodontidae
- Family Polygyridae
- Family Sphincterochilidae
- Family Thysanophoridae
- Family Trissexodontidae
- Family Xanthonychidae

2010 taxonomy

Jörger et al. (2010)[1] analyzed major groups within the Heterobranchia using genetic data and found that Pulmonata as traditionally defined was polyphyletic, for instance some pulmonates were more closely related to Sacoglossa and Acochlidia. They proposed the more inclusive taxon Panpulmonata to unite the clades Siphonarioidea, Sacoglossa, Glacidorboidea, Pyramidelloidea, Amphiboloidea, Hygrophila, Acochlidia and Eupulmonata.[1]

參考文獻

Jörger, Katharina M; Stöger, Isabella; Kano, Yasunori; Fukuda, Hiroshi; Knebelsberger, Thomas; Schrödl, Michael. . BMC Evolutionary Biology. 2010, 10 (1): 323. PMC 3087543. PMID 20973994. doi:10.1186/1471-2148-10-323.
Pek, I.; Vašíček, Z.; Roček, Z.; Hajn., V.; Mikuláš, R. . Olomouc. 1996: 264 pp. ISBN 80-7067-599-3 （捷克语）.
Barnes, Robert D. . Philadelphia, PA: Holt-Saunders International. 1982: 377. ISBN 0-03-056747-5.
Frýda, J.; Hausdorf, B.; Ponder, W.; Valdés, Á.; Warén, A. Bouchet P.; Rocroi J.-P. , 编. . Malacologia: International Journal of Malacology (Hackenheim, Germany / Ann Arbor, USA: ConchBooks). 2005, 47 (1-2): 397 pp. ISBN 3-925919-72-4. ISSN 0076-2997.
Dayrat B. (preview). Zootaxa. 2009, 2068: 1–26 [2016-10-28]. （原始内容存档 (PDF)于2019-07-24）.
吴岷 (编). . . 无脊椎动物第58卷艾纳螺总科第一版. 科学出版社. 2018-06-01: 229-234 [2018-12-27]. ISBN 9787030574831 （中文（简体）‎）.
Wade, C. M.; Mordan, P. B.; Clarke, B. . Proceedings of the Royal Society B: Biological Sciences. 2001, 268 (1465): 413–422. PMC 1088622. PMID 11270439. doi:10.1098/rspb.2000.1372.
Holznagel, W. E.; Colgan, D. J.; Lydeard, C. . Molecular Phylogenetics and Evolution. 2010, 57 (3): 1017–1025. PMID 20920591. doi:10.1016/j.ympev.2010.09.021 （英语）.

外部連結

維基物種中有關肺螺類的數據
. NCBI （英语）.

Pulmonate snails & slugs in captivity

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[Jörger_2010-1] Jörger, Katharina M; Stöger, Isabella; Kano, Yasunori; Fukuda, Hiroshi; Knebelsberger, Thomas; Schrödl, Michael. . BMC Evolutionary Biology. 2010, 10 (1): 323. PMC 3087543. PMID 20973994. doi:10.1186/1471-2148-10-323.

[Pek1996-2] Pek, I.; Vašíček, Z.; Roček, Z.; Hajn., V.; Mikuláš, R. . Olomouc. 1996: 264 pp. ISBN 80-7067-599-3 （捷克语）.

[IZ-3] Barnes, Robert D. . Philadelphia, PA: Holt-Saunders International. 1982: 377. ISBN 0-03-056747-5.

[4] Frýda, J.; Hausdorf, B.; Ponder, W.; Valdés, Á.; Warén, A. Bouchet P.; Rocroi J.-P. , 编. . Malacologia: International Journal of Malacology (Hackenheim, Germany / Ann Arbor, USA: ConchBooks). 2005, 47 (1-2): 397 pp. ISBN 3-925919-72-4. ISSN 0076-2997.

[Dayrat2009-5] Dayrat B. (preview). Zootaxa. 2009, 2068: 1–26 [2016-10-28]. （原始内容存档 (PDF)于2019-07-24）.

[zgdwz_W58-6] 吴岷 (编). . . 无脊椎动物第58卷艾纳螺总科第一版. 科学出版社. 2018-06-01: 229-234 [2018-12-27]. ISBN 9787030574831 （中文（简体）‎）.

[Wade2001-7] Wade, C. M.; Mordan, P. B.; Clarke, B. . Proceedings of the Royal Society B: Biological Sciences. 2001, 268 (1465): 413–422. PMC 1088622. PMID 11270439. doi:10.1098/rspb.2000.1372.

[Holznagel2010-8] Holznagel, W. E.; Colgan, D. J.; Lydeard, C. . Molecular Phylogenetics and Evolution. 2010, 57 (3): 1017–1025. PMID 20920591. doi:10.1016/j.ympev.2010.09.021 （英语）.