布歇特和洛克羅伊的腹足類分類 (2005年)

布歇特和洛克羅伊的腹足類分類 (2005年)(Taxonomy of the Gastropoda (Bouchet & Rocroi, 2005))是一個由巴黎自然史博物館菲利普·布歇特()和讓-皮埃爾·洛克羅伊()於2005年編輯及出版的腹足類軟體動物生物分類系統,以取代過時的腹足纲分类表 (1997年)。所謂腹足類,就是指各種水生或陸生的蝸牛淡水螺海螺蛞蝓。本分類系統刊登於《Malacologia》期刊,包含所有已知的蝸牛與蛞蝓的,以及腹足類的化石。它亦包括古生代地位未定軟體動物,因為這些物種未能分辨到底是腹足綱還是單板綱[1]。本分類系統實際上是由布歇特及一個五人專家組共同編寫;洛克羅伊並沒有參與編寫,而是與布歇特編寫分類的文獻檢索和編輯。

整個分類分為兩部份:第一部份是按照ICZN第四版的命名規則而為各個科-族層級分類的統一命名;而第二部份則是各個分類按其位置而組合的分類樹。整個分類編輯時間歷時二十年。

概說

布歇特和洛克羅伊的系統是首個以支序分類學為主的完整腹足類分類系統,透過從分子系統發生學的研究成果描劃出來。它不再採用傳統林奈分類法總科之間加插的總目亞目等有等級分類,而是採用較自然及統計性的叢集分析而得出的支序演化分支)。在布歇特和洛克羅伊的系統,腹足類生物被劃分為六個主要的支序:

當中,頭三個支序並沒有再包含其他支序,而是直接跳到總科階段。在新生腹足類之下還有另外一個支序。相對來說,異鰓類是綱內最複雜的:單單是裸鰓類就在總科以上有六個支序;而對於其他蝸牛,在總科之上有四個支序。

主要分支

笠形腹足類

笠形腹足類生物在本分類被認為是一個支序,而不是分類單元。這個支序包含了多個科與總科,詳列如下(†表示已滅絕):

  • 笠螺總科 Patelloidea
    • 笠螺科 Patellidae
  • 花笠螺总科 Nacelloidea Thiele, 1891
    • 花笠螺科 Nacellidae Thiele, 1891
  • 蓮花青螺總科 Lottioidea
    • 蓮花青螺科 Lottiidae
    • 青螺科 Acmaeidae Forbes, 1850
      • 青螺亞科 Acmaeinae Forbes, 1850
      • 梳齒笠螺亞科[2] Pectinodontinae Pilsbry, 1891
      • subfamily Rhodopetalinae Lindberg, 1981
    • 无鳃笠螺科 Lepetidae[3]
  • Superfamily Neolepetopsoidea
    • Family Neolepetopsidae
    • † Family Daminilidae
    • † Family Lepetopsidae

與先前旁得和林德伯格的腹足類分類 (1997年)的分類相比較,除了笠形腹足類生物在當時被訂為一個目級分類而不是支序,本類物種其實沒有多大變動。布歇特和洛克羅伊的分類索性跳過旁得和林德伯格分類的亞目,再加上Neolepetopsoidea這個只包含兩個已滅絕科的總科。

古腹足類

古腹足類(Vetigastropoda Salvini-Plawen, 1989)基本上包括了各種笠螺帽貝,覆蓋了原來原始腹足目的大多數物種,計有:

  • 總科地位未定
    • the families: Ataphridae Cossmann, 1915, Crosseolidae Hickman, 2013, Pendromidae Warén, 1991 (synonym: Trachysmatidae Thiele, 1925), † Schizogoniidae
    • the genera: Sahlingia Warén & Bouchet, 2001
  • Superfamily †Amberleyoidea
  • Superfamily †Eotomarioidea
  • 裂螺总科 Fissurelloidea Fleming, 1822:亦作裂裙螺總科,是海螺的一個特化體,其下只有裂螺科一個科。
  • 鲍螺总科 Haliotoidea Rafinesque, 1815
  • Superfamily Lepetelloidea
  • 龙角螺总科 Lepetodriloidea McLean, 1988 (热液孔笠螺)
  • 莫氏螺总科Murchisonioidea Koken, 1889:舊屬莫氏螺目
  • Superfamily Neomphaloidea:belongs in its own independent clade, the Neomphalina, outside the Vetigastropoda [4]
  • 翁戎螺总科 Pleurotomarioidea Swainson, 1840:翁戎螺,Neomphalina的姊妹支序[4]
  • Superfamily †Porcellioidea
  • Superfamily Scissurelloidea
  • 陀螺总科 Seguenzioidea Verrill, 1884
  • 钟螺总科 Trochoidea Rafinesque, 1815
  • 蠑螺總科 Turbinoidea

仁螺總目

舊分類仁螺总目(Cocculiniformia Haszprunar, 1987)原被分為兩大總科:

因此,新的分類實際上只餘下科庫螺總科之下的兩個科。

蜑形類

蜑形類支序包括有原來蜑螺總科(Neritacea)及其下各科,如:蜑螺科(Neritidae)。大致可以分為下列三項:

新生腹足類

新生腹足類物種大致分為以下三大分支:

包括下列各科和總科:

  • 新進腹足類的地位未定物種
    • † Family Plicatusidae
    • † Family Spanionematidae
    • † Family Spirostylidae
  • Superfamily Acteoninoidea
    • † Family Acteoninidae
    • † Family Anozygidae
    • † Family Soleniscidae
  • Superfamily Dendropupoidea
    • † Family Dendropupidae
    • † Family Anthracopupidae
  • Superfamily Paleostyloidea
    • † Family Palaeostylidae
    • † Family Goniasmatidae
    • † Family Pithodeidae
  • Superfamily Peruneloidea
    • † Family Perunelidae
    • † Family Chuchlinidae
    • † Family Imoglobidae
    • † Family Sphaerodomidae
  • Superfamily Pseudomelanioidea
    • † Family Pseudomelaniidae
    • † Family Trajanellidae
  • Superfamily Subulitoidea
    • † Family Subulitidae
    • † Family Ischnoptygmatidae
  • grade Zygopleuroid group - see Changes since 2005
    • † Family Zygopleuridae
    • Family Abyssochrysidae
    • † Family Polygyrinidae
    • † Family Protorculidae
    • Family Provannidae
    • † Family Pseudozygopleuridae

新進腹足類的地位未定物種

新進腹足類之下現時地位未定物種都是化石種,包括下列三個科:

  • † Family Plicatusidae
  • † Family Spanionematidae
  • † Family Spirostylidae

異鰓類

異鰓類是綱內最複雜的:單單是裸鰓類就在總科以上有六個支序;而對於其他蝸牛,在總科之上有四個支序。簡單來說,異鰓類物種包括了原來的後鰓目有肺目的物種,原來的基眼亚目在2005年的分類裡成為了有肺目之下的一個非正式群組裡,當中包括喜濕螺類;而原來的有肺目跟囊舌類等其他物種又組成泛有肺類。其餘沒有被歸類的物種,皆籠統的被分類到下異鰓類這一個非正式群組裡。

系統分類

在下列分類中,只有被歸為總科級和科級的分類才會被縮排。不分的支序不會被縮排,但這些名字的字體會改變。有關支序間的從屬關係可參看對上一節的列表。

古生代地位未定軟體動物

古生代地位未定軟體動物Paleozoic molluscs of uncertain systematic position)都只有化石種,大致可以分為下列三個種類:

未能分辨到底是腹足綱還是單板綱物種

以下分類元及物種未能分辨到底是腹足綱還是單板綱Monoplacophora

  • 總科地位未定
    • Khairkhaniidae
    • Ladamarekiidae
    • Metoptomatidae
    • Patelliconidae
    • Protoconchoididae
  • Archinacelloidea
    • Archinacellidae
    • Archaeopragidae
  • Pelagielloidea
    • Pelagiellidae
    • Aldanellidae
  • Scenelloidea
    • Scenellidae
    • Coreospiridae
    • Igarkiellidae
  • Yochelcionelloidea
    • Yochelcionellidae
    • Stenothecidae
    • Trenellidae

With isostrophically coiled shells of uncertain position (Gastropoda or Monoplacophora)

    • Bucanellidae
    • Bucaniidae
    • Euphemitidae
    • Pterothecidae
    • Sinuitidae
    • Tremanotidae
    • Tropidodiscidae

With anisostrophically coiled shells of uncertain position (Gastropoda?)

  • Euomphaloidea
    • Euomphalidae
    • Helicotomidae
    • Lesueurillidae
    • Omphalocirridae
    • Omphalotrochidae
  • Macluritoidea
    • Macluritidae

已確認為腹足綱基群的分類元

(只包括化石種分類元)

  • 總科分類未定
    • Family Anomphalidae † :2017年分類不再存在。
    • Family Codonocheilidae S. A. Miller, 1889
    • Family Crassimarginatidae Frýda, Blodgett & Lenz, 2002
    • Family Holopeidae † :2017年分類不再存在。
    • Family Isospiridae Wangberg-Eriksson, 1964
    • Family Opisthonematidae Yu, 1976 †:後來在2017年的分類為Yuopisthonematidae Nützel, 2017 † 所取代,因為其模式屬成為了異名。
    • 擬蠑螺科 Paraturbinidae Cossmann, 1916
    • Family Planitrochidae † :2017年分類不再存在。
    • Family Pragoserpulinidae Frýda, 1998
    • Family Pseudophoridae † :2017年分類不再存在。
    • Family Raphistomatidae Koken, 1896
    • Family Rhytidopilidae Starobogatov, 1976
    • Family Scoliostomatidae Frýda, Blodgett & Lenz, 2002
    • Family Sinuopeidae Wenz, 1938
  • Superfamily Clisospiroidea S. A. Miller, 1889
    • Clisospiridae
    • Onychochilidae
  • 曲线螺总科 Loxonematoidea Koken, 1889
    • 曲线螺科 Loxonematidae Koken, 1889
    • Palaeozygopleuridae
  • Superfamily Ophiletoidea Koken, 1907
    • Ophiletidae
  • Superfamily Straparollinoidea P. J. Wagner, 2002 †:後來在2017年的分類被認為是Euomphaloidea White, 1877 †的異名。
    • Straparollinidae
  • Superfamily Trochonematoidea Zittel, 1895
    • Trochonematidae
    • Lophospiridae

笠形腹足類

笠形腹足類支序(Patellogastropoda)在本分類中包括下列四個總科:

  • 笠螺總科 Patelloidea
    • 笠螺科 Patellidae
  • 花笠螺总科 Nacelloidea Thiele, 1891
    • 花笠螺科 Nacellidae Thiele, 1891
  • 蓮花青螺總科 Lottioidea
    • 蓮花青螺科 Lottiidae
    • 青螺科 Acmaeidae Forbes, 1850
      • 青螺亞科 Acmaeinae Forbes, 1850
      • 梳齒笠螺亞科[2] Pectinodontinae Pilsbry, 1891
      • subfamily Rhodopetalinae Lindberg, 1981
    • 无鳃笠螺科 Lepetidae[3]
  • Superfamily Neolepetopsoidea
    • Family Neolepetopsidae
    • † Family Daminilidae
    • † Family Lepetopsidae

這四個總科之下的分類元在往後的文獻曾作多番修訂,至今仍未有定案。詳見各分支的條目內容。

古腹足類支序

古腹足類支序(Vetigastropoda Salvini-Plawen, 1989)的成員計有:

  • Not assigned to a superfamily:
    • Family Ataphridae
    • Family Pendromidae
    • † Family Schizogoniidae
  • Superfamily †Amberleyoidea
    • † Family Amberleyidae
    • † Family Nododelphinulidae
  • Superfamily †Eotomarioidea
    • † Family Eotomariidae
    • † Family Gosseletinidae
    • † Family Luciellidae
    • † Family Phanerotrematidae
  • Superfamily Fissurelloidea
  • 鮑總科 Haliotoidea
  • Superfamily Lepetelloidea
    • Family Lepetellidae
    • Family Addisoniidae
    • Family Bathyphytophilidae
    • Family Caymanabyssiidae
    • Family Cocculinellidae
    • Family Osteopeltidae
    • Family Pseudococculinidae
    • Family Pyropeltidae
  • Superfamily Lepetodriloidea
    • Family Lepetodrilidae
    • Family Clypeosectidae
    • Family Sutilizonidae
  • Superfamily †Murchisonioidea
    • † Family Murchisoniidae
    • † Family Cheeneetnukiidae
    • † Family Hormotomidae
  • Superfamily Neomphaloidea: subsequent research has placed this superfamily in its own clade, the Neomphalina, basal to the Vetigastropoda [5]
    • Family Neomphalidae
    • Family Melanodrymiidae
    • Family Peltospiridae
  • Superfamily Pleurotomarioidea
    • Family Pleurotomariidae
    • † Family Catantostomatidae
    • † Family Kittlidiscidae
    • † Family Phymatopleuridae
    • † Family Polytremariidae
    • † Family Portlockiellidae
    • † Family Rhaphischismatidae
    • † Family Trochotomidae
    • † Family Zygitidae
  • Superfamily †Porcellioidea
    • † Family Porcelliidae
    • † Family Cirridae
    • † Family Discohelicidae
    • † Family Pavlodiscidae
  • Superfamily Scissurelloidea
    • Family Scissurellidae
    • Family Anatomidae
  • Superfamily Seguenzioidea
    • Family Seguenziidae
    • Family Chilodontidae
    • † Family Eucyclidae
    • † Family Laubellidae
  • Superfamily Trochoidea
    • Family Trochidae
    • Family Calliostomatidae
    • † Family Elasmonematidae
    • † Family Eucochlidae
    • † Family Microdomatidae
    • † Family Proconulidae
    • Family Solariellidae
    • † Family Tychobraheidae
    • † Family Velainellidae
  • Superfamily Turbinoidea

Clade Cocculiniformia

  • Superfamily Cocculinoidea
    • Family Cocculinidae
    • Family Bathysciadiidae

蜑形類支序

蜑形類支序的三大分支如下:

地位未定古生代蜑形類物種

  • 總科地位未定
    • Family Craspedostomatidae Wenz, 1938
      • Subfamily Craspedostomatinae Wenz, 1938
      • Subfamily Bucanospirinae Wenz, 1938[6]
    • † Family Pragoscutulidae
  • Superfamily Nerrhenoidea
    • † Family Nerrhenidae
  • Superfamily Oriostomatoidea
    • † Family Oriostomatidae
    • † Family Tubinidae
  • 古輪螺總科 Palaeotrochoidea
  • Superfamily Platyceratoidea
    • † Family Platyceratidae

clade Cyrtoneritimorpha

clade Cyrtoneritimorpha

  • 總科地位未定
    • † Family Orthonychiidae
    • † Family Vltaviellidae

Cycloneritimorpha支序

  • 樹螺總科Helicinoidea
    • 樹螺科Helicinidae
    • † Family Dawsonellidae
    • † Family Deaniridae
    • Family Neritiliidae
    • Family Proserpinellidae
    • Family Proserpinidae
  • Superfamily Hydrocenoidea
    • Family Hydrocenidae
  • 蜑螺總科Neritoidea
    • 蜑螺科Neritidae
    • Family Phenacolepadidae
    • † Family Pileolidae
  • 珍珠蜑螺總科Neritopsoidea
    • 珍珠蜑螺科Neritopsidae
    • Family Neritopsidae
    • † Family Cortinellidae
    • † Family Delphinulopsidae
    • † Family Plagiothyridae
    • † Family Pseudorthonychiidae
    • Family Titiscaniidae
  • Superfamily Symmetrocapuloidea
    • † Family Symmetrocapulidae

新生腹足類支序

新生腹足類支序(Caenogastropoda)除了主扭舌類吸螺類高腹足類三大分支以外,還包括其系統分類地位未定的分類元,分別如下:

  • 新生腹足類支序之下系統分類地位未定的科:
    • † Family Plicatusidae
    • † Family Spanionematidae
    • † Family Spirostylidae
  • Superfamily Acteoninoidea
    • † Family Acteoninidae
    • † Family Anozygidae
    • † Family Soleniscidae
  • Superfamily Dendropupoidea
    • † Family Dendropupidae
    • † Family Anthracopupidae
  • Superfamily Paleostyloidea
    • † Family Palaeostylidae
    • † Family Goniasmatidae
    • † Family Pithodeidae
  • Superfamily Peruneloidea
    • † Family Perunelidae
    • † Family Chuchlinidae
    • † Family Imoglobidae
    • † Family Sphaerodomidae
  • Superfamily Pseudomelanioidea
    • † Family Pseudomelaniidae
    • † Family Trajanellidae
  • Superfamily Subulitoidea
    • † Family Subulitidae
    • † Family Ischnoptygmatidae
  • grade Zygopleuroid group - see Changes since 2005
    • † Family Zygopleuridae
    • Family Abyssochrysidae
    • † Family Polygyrinidae
    • † Family Protorculidae
    • Family Provannidae
    • † Family Pseudozygopleuridae

主扭舌類非正式群組

主扭舌類非正式群組(Informal group Architaenioglossa)包括下列各個總科與科:

此外,根據其他文獻,深水螺總科被認為跟本群組的

吸螺類支序

吸螺類Sorbeoconcha)原為吸螺目,亦作吸腔目,由三個亞目組成:

以下為本支序之下各總科與科:

  • 總科地位未定
    • † Family Acanthonematidae
    • † Family Canterburyellidae
    • † Family Prisciphoridae
  • 蟹守螺總科 Cerithioidea
    • 蟹守螺科 Cerithiidae
    • Family Batillariidae
    • † Family Brachytrematidae
    • † Family Cassiopidae
    • Family Dialidae
    • Family Diastomatidae
    • † Family Eustomatidae
    • † Family Ladinulidae
    • † Family Lanascalidae
    • Family Litiopidae
    • † Family Maoraxidae
    • Family Melanopsidae
    • † Family Metacerithiidae
    • Family Modulidae
    • Family Pachychilidae
    • Family Paludomidae
    • Family Planaxidae
    • Family Pleuroceridae
    • † Family Popenellidae
    • Family Potamididae
    • † Family Procerithiidae - If the genus Argyropeza is placed in the Procerithiidae, then this family is not all fossil.
    • † Family Prostyliferidae
    • † Family Propupaspiridae
    • Family Scaliolidae
    • Family Siliquariidae
    • † Family Terebrellidae - not a valid name
    • Family Thiaridae
    • 錐螺科 Turritellidae
  • 鐘形蟹守螺總科 Campaniloidea

高腹足類支序

高腹足類Hypsogastropoda)包括有 玉黍螺類支序(Littorinimorpha)、新腹足類支序 (Neogastropoda)及翼舌類非正式群組(Ptenoglossa),還有以下不屬於任何總科的四個化石科:

  • † Family Coelostylinidae
  • † Family Maturifusidae
  • † Family Pommerozygiidae
  • † Family Settsassiidae

玉黍螺類支序

玉黍螺類支序Littorinimorpha)包括下列各個總科與科:

翼舌類非正式群組

翼舌類Ptenoglossa)這個非正式的群組是一個並系群,包括下列三個總科:

新腹足類支序

新腹足類支序(Neogastropoda)包括以下各個總科:

  • Unassigned to a superfamily
    • † Family Johnwyattiidae
    • † Family Perissityidae
    • † Family Sarganidae
    • † Family Speightiidae
    • † Family Taiomidae
    • † Family Weeksiidae
  • 峨螺總科 Buccinidae Rafinesque, 1815
  • Superfamily Muricoidea
    • Family Muricidae
    • Family Babyloniidae
    • Family Costellariidae
    • Family Cystiscidae
    • Family Harpidae
    • Family Marginellidae
    • Family Mitridae
    • † Family Pholidotomidae
    • Family Pleioptygmatidae
    • Family Strepsiduridae
    • Family Turbinellidae
    • Family Volutidae
    • Family Volutomitridae
  • Superfamily Olivoidea
    • Family Olividae
    • Family Olivellidae
  • Superfamily Pseudolivoidea
    • Family Pseudolividae
    • Family Ptychatractidae
  • Superfamily Conoidea
    • Family Conidae
    • Family Clavatulidae
    • Family Drilliidae
    • Family Pseudomelatomidae
    • Family Strictispiridae
    • Family Terebridae
    • Family Turridae
  • Superfamily Cancellarioidea
    • Family Cancellariidae

異鰓類支序

異鰓類支序(Heterobranchia)大致可以分成以下三個非正式群組:

  • 異鰓類(Heterobranchia)
  • 後鰓類(Opisthobranchia)及
  • 有肺類(Pulmonata)

下異鰓類非正式群組

下異鰓類非正式群組("Lower Heterobranchia";= Allogastropoda

  • Unassigned to a superfamily
    • Family Cimidae
    • † Family Dolomitellidae
    • † Family Heterosubulitidae
    • † Family Kuskokwimiidae
    • † Family Misurinellidae
    • Family Orbitestellidae
    • Family Tjaernoeiidae
    • Family Xylodisculidae
  • Superfamily Acteonoidea
  • Superfamily Architectonicoidea
  • Superfamily Glacidorboidea
  • Superfamily Mathildoidea
    • Family Mathildidae
    • † Family Ampezzanildidae
    • † Family Anoptychiidae
    • † Family Gordenellidae
    • † Family Tofanellidae
    • † Family Trachoecidae
  • Superfamily Nerineoidea
    • † Family Nerineidae
    • † Family Ceritellidae
    • † Family Nerinellidae
  • Superfamily Omalogyroidea
    • Family Omalogyridae
    • † Family Studraxidae
  • Superfamily Pyramidelloidea
    • Family Pyramidellidae
    • Family Amathinidae
    • † Family Heteroneritidae
    • Family Murchisonellidae
  • 露齒螺總科 Ringiculoidea
    • 露齒螺科 Ringiculidae
  • Superfamily Rissoelloidea
    • Family Rissoellidae
  • Superfamily Streptacidoidea
    • † Family Streptacididae
    • † Family Cassianebalidae
  • Superfamily Valvatoidea
    • Family Valvatidae
    • Family Cornirostridae
    • Family Hyalogyrinidae
    • † Family Provalvatidae

後鰓類

後鰓類非正式群組(Opisthobranchia)源自原來的後鰓目,包括以下各個分類:

頭楯目

頭楯目Cephalaspidea)包括以下五個總科:

有殼翼足類

有殼翼足類支序Thecosomata)包括以下各個科與總科:

  • Superfamily Cavolinioidea
    • Family Cavoliniidae
    • Family Limacinidae
    • † Family Sphaerocinidae
  • Superfamily Cymbulioidea
    • Family Cymbuliidae
    • Family Desmopteridae
    • Family Peraclidae

裸體翼足類

裸體翼足類支序(Gymnosomata)包括以下兩個總科:

  • Superfamily Clionoidea
    • Family Clionidae
    • Family Cliopsidae
    • Family Notobranchaeidae
    • Family Pneumodermatidae
  • Superfamily Hydromyloidea
    • Family Hydromylidae
    • Family Laginiopsidae

Clade Aplysiomorpha (= Anaspidea)

Group Acochlidiacea

  • Superfamily Acochlidioidea
    • Family Acochlidiidae
  • Superfamily Hedylopsoidea
    • Family Hedylopsidae
    • Family Ganitidae
    • Family Livorniellidae
    • Family Minicheviellidae
    • Family Parhedylidae
    • Family Tantulidae
  • Superfamily Palliohedyloidea
    • Family Palliohedylidae
  • Superfamily Strubellioidea
    • Family Strubelliidae
    • Family Pseudunelidae

囊舌類支序

囊舌類支序Sacoglossa)包括下列兩個亞支序:

長足亞支序

長足亞支序(Oxynoacea)皆為有殼物種[11],成員如下:

  • 长足总科 Oxynooidea Stoliczka, 1868 (1847):本總科物種。
    • 长足科 Oxynoidae Stoliczka, 1868 (1847)
    • 珠綠螺科 Juliidae E. A. Smith, 1885
    • 圆卷螺科 Volvatellidae Pilsbry, 1895

海天牛亞支序

海天牛亞支序(Placobranchacea)主要成員皆為沒有殼的物種,詳細如下:

  • 海天牛總科 Placobranchoidea Gray, 1840
    • 海天牛科 Placobranchidae Gray, 1840
    • Family Boselliidae
    • Family Platyhedylidae Salvini-Plawen, 1973
  • 柱状總科 Limapontioidea Gray, 1847
    • 柱状科 Limapontiidae Gray, 1847:舊作棍螺科 Stiligeridae[12]
    • Family Caliphyllidae
    • 荷叶鳃科 Hermaeidae H. Adams & A. Adams, 1854

Group Cylindrobullida

傘螺類支序

傘螺類支序(Umbraculida)包括由下列兩個科組成的傘螺總科

  • 傘螺總科 Umbraculoidea
    • 傘螺科 Umbraculidae
    • Family Tylodinidae

裸側類

裸側類支序(Nudipleura)由側鰓類亞支序裸鰓類亞支序

側鰓類亞支序

側鰓類亞支序(Pleurobranchomorpha)只有一個總科:

裸鰓類亞支序

裸鰓類亞支序(Nudibranchia)大致可以分為下列三個部份:

有角鰓類支序

有角鰓類支序(Euctenidiacea ,即Holohepatica) Contains the subclades Gnathodoridacea and 海蛞蝓亞支序

Subclade Gnathodoridacea
  • Superfamily Bathydoridoidea
    • Family Bathydorididae
海蛞蝓亞支序
  • Superfamily Doridoidea
    • Family Dorididae
    • Family Actinocyclidae
    • Family Chromodorididae
    • Family Discodorididae
  • 葉海牛總科 Phyllidioidea Rafinesque, 1814
    • 葉海牛科 Phyllidiidae Rafinesque, 1814
    • 枝鰓海麒麟科 Dendrodorididae O'Donoghue, 1924 (1864)
    • Family Mandeliidae Valdés & Gosliner, 1999
  • Superfamily Onchidoridoidea
    • Family Onchidorididae
    • Family Corambidae
    • Family Goniodorididae
  • 多角海蛞蝓總科 Polyceroidea (= Phanerobranchiata Non Suctoria)
    • 多角海蛞蝓科 Polyceridae
    • Family Aegiretidae - Aegiretidae is incorrect subsequent spelling of this family in Bouchet & Rocroi (2005). Correct is Aegiridae.
    • Family Gymnodorididae
    • Family Hexabranchidae
    • Family Okadaiidae
無角鰓類支序

無角鰓類支序(Dexiarchia,即Actenidiacea)包括Pseudoeuctenidiacea and Cladobranchia兩個支序:

Clade Pseudoeuctenidiacea ( = Doridoxida)
  • Superfamily Doridoxoidea
    • Family Doridoxidae
Clade Cladobranchia ( = Cladohepatica)

Contains the subclades Euarminida, Dendronotida and Aeolidida

  • Not assigned to a superfamily
    • Family Charcotiidae
    • Family Dironidae
    • Family Dotidae
    • Family Embletoniidae
    • Family Goniaeolididae
    • Family Heroidae
    • Family Madrellidae
    • Family Pinufiidae
    • Family Proctonotidae
Subclade Euarminida
  • Superfamily Arminoidea
    • Family Arminidae
    • Family Doridomorphidae
枝鰓類

枝鰓類Dendronotida[13]包括下列各科:

蓑海牛類

蓑海牛類Aeolidida)亞支序包括下列各分類單元:

  • Superfamily Flabellinoidea ( = Pleuroprocta)
    • Family Flabellinidae
    • Family Notaeolidiidae
  • Superfamily Fionoidea
    • Family Fionidae
    • Family Calmidae
    • Family Eubranchidae
    • Family Pseudovermidae
    • Family Tergipedidae
  • 蓑海牛总科 Aeolidioidea

有肺類支序

有肺類支序(Pulmonata)包括非正式群組基眼類真有肺類支序

非正式群組基眼類

非正式群組基眼類Basommatophora)除包括高腹足類支序以外,還包括下列兩個總科:

  • 兩棲螺總科 Amphiboloidea
  • Superfamily Siphonarioidea
    • Family Siphonariidae
    • † Family Acroreiidae
喜濕螺類支序

喜濕螺類支序(Hygrophila)包括以下各個總科:

  • 斜顶螺总科 Acroloxoidea Thiele, 1931
    • 斜顶螺科 Acroloxidae Thiele, 1931
  • Superfamily Chilinoidea Dall, 1870
    • Family Chilinidae Dall, 1870
    • Family Latiidae Hutton, 1882
  • 椎实螺总科(Lymnaeoidea Rafinesque, 1815)
    • 椎实螺科Lymnaeidae Rafinesque, 1815)
  • 扁蜷總科Planorboidea Rafinesque, 1815)

真有肺類支序

真有肺類支序(Eupulmonata)除包括收眼類支序柄眼類支序兩個支序以外,還包括下列各總科與科:

收眼類支序

收眼類支序(Systellommatophora=Gymnomorpha)之下有兩總科[14],分別為:

柄眼類支序

柄眼類支序(Stylommatophora)包括下列兩個亞支序與一個非正式群組:

板顎類亞支序

板顎類亞支序(Subclade Elasmognatha)包括:

  • 琥珀螺超科()
    • 琥珀螺科()
  • 无盔蛞蝓超科()
    • 无盔蛞蝓科()
直尿道類支序

直尿道類支序(Orthurethra)包括下列各總科與科:

  • 帕图螺超科()
    • 帕图螺科()
  • 小玛瑙螺超科()
    • 小玛瑙螺科()
  • 槲果螺超科()
  • 蛹蜗牛超科()
    • 虹蛹螺科()
    • 粒蛹螺科()
    • (已灭绝)
    • 蛹状螺科()
    • 球果螺科()
    • 瓦娄蜗牛科()
    • 旋螺科()
  • 艾纳螺超科()
彎尿道類非正式群組

彎尿道類非正式群組(Sigmurethra)包括下列各分類單元:

  • 烟管蜗牛总科 Clausilioidea Gray, 1855  
  • 彩带螺超科()
    • 彩带螺科()
    • 蜂窝蜗牛科()
    • (已灭绝)
    • 尾刺螺科()
  • 非洲大蝸牛總科 Achatinoidea Swainson, 1840
  • 爱丽蛞蝓超科()
    • 爱丽蛞蝓科()
  • 小壳螺超科()
    • 小壳螺科()
    • 橡子螺科()
    • 旋轴螺科()
  • 扭轴蜗牛超科()
  • 彩釉蜗牛超科()
    • 彩釉蜗牛科()
    • 斗蓬螺科()
    • 单孔螺科()
  • 红蜗牛超科()
    • 针状螺科()
    • 硬壳螺科()
    • 羚管螺科()
    • 玫瑰口蜗牛科()
  • 圈螺超科()
  • 粟蜗牛超科()
    • 粟蜗牛科()
    • (已灭绝)
    • 茶色蜗牛科()
    • 内齿螺科()
    • 山地蜗牛科()
    • 盾状螺科()
  • 袋螺超科()
    • 袋螺科()
  • 旋蜗牛超科()
    • 大蜗牛科()
    • 巴蜗牛科()
    • 坚齿螺科()
    • 扁雕蜗牛科()
    • 嗜石螺科()
    • 肋齿螺科()
    • 多圆螺科()
  • 腹齒螺總科 [15]
  • 小盾螺超科()
    • 小盾螺科()
    • 小蛞蝓科()
    • 三角外套蛞蝓科()
  • 带螺超科()
    • 带螺科()
  • 鳖甲蜗牛超科()
  • 背盾蛞蝓超科()
  • 欧洲蛞蝓超科()
    • 阿勇蛞蝓科()
    • 高山蛞蝓科()
    • 欧洲蛞蝓科()
    • 嗜粘液蛞蝓科()
  • Superfamily Aillyoidea
    • Family Aillyidae
  • Superfamily Testacelloidea
    • Family Testacellidae
    • Family Oleacinidae
    • Family Spiraxidae
  • Superfamily Papillodermatoidea
    • Family Papillodermatidae
  • Superfamily Streptaxoidea
  • Superfamily Rhytidoidea
    • Family Rhytididae
    • Family Chlamydephoridae
    • Family Haplotrematidae
    • Family Scolodontidae
  • Superfamily Acavoidea
    • Family Acavidae
    • Family Caryodidae
    • Family Dorcasiidae
    • Family Macrocyclidae
    • Family Megomphicidae
    • Family Strophocheilidae
  • Superfamily Plectopyloidea
    • Family Plectopylidae
    • Family Corillidae
    • Family Sculptariidae
  • Superfamily Punctoidea
    • Family Punctidae
    • † Family Anastomopsidae
    • Family Charopidae
    • Family Cystopeltidae
    • Family Discidae
    • Family Endodontidae
    • Family Helicodiscidae
    • Family Oreohelicidae
    • Family Thyrophorellidae
  • Superfamily Sagdoidea
    • Family Sagdidae
limacoid clade
  • Superfamily Staffordioidea
    • Family Staffordiidae
  • Superfamily Dyakioidea
    • Family Dyakiidae
  • Superfamily Gastrodontoidea
    • Family Gastrodontidae
    • Family Chronidae
    • Family Euconulidae
    • Family Oxychilidae
    • Family Pristilomatidae
    • Family Trochomorphidae
    • Fossil taxa that probably belong to the superfamily Gastrodontoidea:
      • Subfamily † Archaeozonitinae
      • Subfamily † Grandipatulinae
      • Subfamily † Palaeoxestininae
  • Superfamily Parmacelloidea
    • Family Parmacellidae
    • Family Milacidae
    • Family Trigonochlamydidae
  • Superfamily Zonitoidea
  • Superfamily Helicarionoidea
  • Superfamily Limacoidea
Informal group Sigmurethra continued

Two other superfamilies are part of the clade Sigmurethra, but they are not in the limacoid clade:

  • Superfamily Arionoidea
    • Family Arionidae
    • Family Anadenidae
    • Family Ariolimacidae
    • Family Binneyidae
    • Family Oopeltidae
    • Family Philomycidae
  • Superfamily Helicoidea
    • Family Helicidae
    • Family Bradybaenidae
    • Family Camaenidae
    • Family Cepolidae
    • Family Cochlicellidae
    • Family Elonidae
    • Family Epiphragmophoridae
    • Family Halolimnohelicidae
    • Family Helicodontidae
    • Family Helminthoglyptidae
    • Family Humboldtianidae
    • Family Hygromiidae
    • Family Monadeniidae
    • Family Pleurodontidae
    • Family Polygyridae
    • Family Sphincterochilidae
    • Family Thysanophoridae
    • Family Trissexodontidae
    • Family Xanthonychidae

參考文獻

  1. Frýda J., Hausdorf B., Ponder W., Valdés Á. & Warén A. . Malacologia: International Journal of Malacology (Hackenheim, Germany: ConchBooks). 2005, 47 (1-2): 397. ISBN 3-925919-72-4. ISSN 0076-2997. 已忽略未知参数|first-editor2= (帮助); 已忽略未知参数|last-editor2= (帮助); 已忽略未知参数|last-editor1= (帮助); 已忽略未知参数|first-editor1= (帮助)
  2. Stephanie W. Aktipis & Gonzalo Giribet (2010). "A phylogeny of Vetigastropoda and other “archaeogastropods”: re-organizing old gastropod clades". Invertebrate Biology 129(3): 220-240 页面存档备份,存于 doi:10.1111/j.1744-7410.2010.00198.x.
  3. WoRMS: Neomphalina
  4. WoRMS. . World Register of Marine Species. [2019-05-18].
  5. . 2011-07-13 [2017-08-17] (中文(繁體)‎).
  6. Dayrat B. (preview). Zootaxa. 2009, 2068: 1–26.
  • Guido T. Poppe; Sheila P. Tagaro. (PDF). Visaya. 2006-02-23 [2018-05-13]. (原始内容 (PDF)存档于2017年7月29日) (英语).

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